Crossword clues for the word: RUGAE The word RUGAE has appeared as an answer to at least 4 crossword clues on different crossword puzzles. Crossword clues for RUGAE Showing 1 to 4 of 4 results Clue(s) "Wrinkles, in botany." "Wrinkles, in zoology." "Wrinkles, to botanists" Wrinkles on Cato's brow